| Andrew William Oliver |
205,882,352,941 |
£319,117,647 |
| TD Direct Investing Nominees |
94,495,144,706 |
£146,467,474 |
| HSDL Nominees Limited |
81,968,497,647 |
£127,051,171 |
| David Lenigas |
79,411,764,706 |
£123,088,235 |
| Greenhair Services Limited |
73,235,294,118 |
£113,514,706 |
| Academy Minerals Limited |
70,588,235,294 |
£109,411,765 |
| Kavi Dhana |
44,021,067,059 |
£68,232,654 |
| IP2IPO Limited (together with Techtran Group Limited) |
34,418,058,824 |
£53,347,991 |
| XCAP Nominees Limited |
19,824,847,059 |
£30,728,513 |
| IP Venture Fund Limited |
16,302,105,882 |
£25,268,264 |
| Mr Mark V Houghton-Berry |
12,335,376,471 |
£19,119,834 |
| The University of Leeds |
9,552,264,706 |
£14,806,010 |
| The Wellcome Trust Limited |
6,107,858,824 |
£9,467,181 |
| Adrian Stiff |
37,000,000 |
£57,350 |
| Hargreaves Lansdown |
27,752,212 |
£43,016 |
| JIM Nominees Limited |
27,116,707 |
£42,031 |
| Pershing Nominees |
26,006,789 |
£40,310 |
| Hargreaves Lansdown (Nominees) Limited Des 15942 |
25,609,943 |
£39,695 |
| Chris Akers |
25,000,000 |
£38,750 |
| Vidacos Nominees Limited |
20,709,150 |
£32,099 |
| The Bank OF New York (Nominees) Limited Des 672938 |
19,705,371 |
£30,543 |
| Oberon Investments |
19,705,371 |
£30,543 |
| Barclays Direct Investing Nominees Limited |
19,358,393 |
£30,006 |
| Winterflood Securities Limited |
18,278,266 |
£28,331 |
| Donald Strang |
17,820,211 |
£27,621 |
| Interactive Investor Services Nominees Limited |
17,219,649 |
£26,690 |
| ABN Amro Bank |
13,801,956 |
£21,393 |
| Hargreaves Lansdown (Nominees) Limited Des VRA |
12,572,701 |
£19,488 |
| David Graham Brown |
12,100,000 |
£18,755 |
| Sebastian Marr |
8,180,672 |
£12,680 |