| Gresham House Asset Management Ltd |
3,884,684 |
£55,162,513 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£46,556,148 |
| Octopus Investments Nominees Limited |
2,335,294 |
£33,161,175 |
| Liontrust Investment Services Limited |
1,743,782 |
£24,761,704 |
| Aberforth Partners LLP |
1,606,852 |
£22,817,298 |
| Liontrust Investment Partners LLP |
1,549,398 |
£22,001,452 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£15,790,613 |
| Brooks Macdonald Asset Management |
1,105,946 |
£15,704,433 |
| Canaccord Genuity Group Inc |
1,021,288 |
£14,502,290 |
| Jupiter Fund Management plc |
938,000 |
£13,319,600 |
| Invesco Asset Management |
817,151 |
£11,603,544 |
| Invesco |
817,151 |
£11,603,544 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£11,490,640 |
| The Wellcome Trust Limited |
805,370 |
£11,436,254 |
| FIDELITY INTERNATIONAL |
786,452 |
£11,167,618 |
| Hargreave Hale & Co |
784,942 |
£11,146,176 |
| Canaccord Genuity Group Inc |
780,008 |
£11,076,114 |
| Artemis Investment Management Ltd |
729,575 |
£10,359,965 |
| Norges Bank |
683,030 |
£9,699,026 |
| Charles Stanley |
659,277 |
£9,361,733 |
| Jonathan M Gumpel |
642,460 |
£9,122,932 |
| Brewin Dolphin Limited |
622,157 |
£8,834,629 |
| Chelverton Asset Management |
609,008 |
£8,647,914 |
| JTC plc |
603,261 |
£8,566,306 |
| Amati Global Investors |
572,459 |
£8,128,918 |
| Rathbones Investment Management |
488,191 |
£6,932,312 |
| Kames Capital |
487,376 |
£6,920,739 |
| Spearpoint Holdings Limited |
428,227 |
£6,080,823 |