| Gresham House Asset Management Ltd |
3,884,684 |
£60,795,305 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£51,310,121 |
| Octopus Investments Nominees Limited |
2,335,294 |
£36,547,351 |
| Aberforth Partners LLP |
1,787,188 |
£27,969,492 |
| Liontrust Investment Services Limited |
1,743,782 |
£27,290,188 |
| Liontrust Investment Partners LLP |
1,549,398 |
£24,248,079 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£17,403,035 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,308,055 |
| Canaccord Genuity Group Inc |
1,021,288 |
£15,983,157 |
| Jupiter Fund Management plc |
938,000 |
£14,679,700 |
| Invesco Asset Management |
817,151 |
£12,788,413 |
| Invesco |
817,151 |
£12,788,413 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£12,663,980 |
| The Wellcome Trust Limited |
805,370 |
£12,604,040 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,307,974 |
| Hargreave Hale & Co |
784,942 |
£12,284,342 |
| Canaccord Genuity Group Inc |
780,008 |
£12,207,125 |
| Artemis Investment Management Ltd |
729,575 |
£11,417,849 |
| Norges Bank |
683,030 |
£10,689,420 |
| Charles Stanley |
659,277 |
£10,317,685 |
| Jonathan M Gumpel |
642,460 |
£10,054,499 |
| Brewin Dolphin Limited |
622,157 |
£9,736,757 |
| Chelverton Asset Management |
609,008 |
£9,530,975 |
| JTC plc |
603,261 |
£9,441,035 |
| Amati Global Investors |
572,459 |
£8,958,983 |
| Rathbones Investment Management |
488,191 |
£7,640,189 |
| Kames Capital |
487,376 |
£7,627,434 |
| Spearpoint Holdings Limited |
428,227 |
£6,701,752 |