| Gresham House Asset Management Ltd |
3,884,684 |
£52,831,702 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£44,588,987 |
| Octopus Investments Nominees Limited |
2,335,294 |
£31,759,998 |
| Liontrust Investment Services Limited |
2,215,411 |
£30,129,590 |
| Aberforth Partners LLP |
1,606,852 |
£21,853,187 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£15,123,404 |
| Brooks Macdonald Asset Management |
1,105,946 |
£15,040,866 |
| Canaccord Genuity Group Inc |
1,021,288 |
£13,889,517 |
| Jupiter Fund Management plc |
938,000 |
£12,756,800 |
| Invesco Asset Management |
817,151 |
£11,113,254 |
| Invesco |
817,151 |
£11,113,254 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£11,005,120 |
| The Wellcome Trust Limited |
805,370 |
£10,953,032 |
| FIDELITY INTERNATIONAL |
786,452 |
£10,695,747 |
| Hargreave Hale & Co |
784,942 |
£10,675,211 |
| Canaccord Genuity Group Inc |
780,008 |
£10,608,109 |
| Artemis Investment Management Ltd |
729,575 |
£9,922,220 |
| Norges Bank |
683,030 |
£9,289,208 |
| Charles Stanley |
659,277 |
£8,966,167 |
| Jonathan M Gumpel |
642,460 |
£8,737,456 |
| Brewin Dolphin Limited |
622,157 |
£8,461,335 |
| Chelverton Asset Management |
609,008 |
£8,282,509 |
| JTC plc |
603,261 |
£8,204,350 |
| Amati Global Investors |
572,459 |
£7,785,442 |
| Rathbones Investment Management |
488,191 |
£6,639,398 |
| Kames Capital |
487,376 |
£6,628,314 |
| Spearpoint Holdings Limited |
428,227 |
£5,823,887 |