| Gresham House Asset Management Ltd |
3,884,684 |
£50,112,424 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£42,293,966 |
| Octopus Investments Nominees Limited |
2,335,294 |
£30,125,293 |
| Liontrust Investment Services Limited |
1,743,782 |
£22,494,788 |
| Aberforth Partners LLP |
1,606,852 |
£20,728,391 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£14,344,994 |
| Brooks Macdonald Asset Management |
1,105,946 |
£14,266,703 |
| Canaccord Genuity Group Inc |
1,021,288 |
£13,174,615 |
| Jupiter Fund Management plc |
938,000 |
£12,100,200 |
| Invesco Asset Management |
817,151 |
£10,541,248 |
| Invesco |
817,151 |
£10,541,248 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£10,438,680 |
| The Wellcome Trust Limited |
805,370 |
£10,389,273 |
| FIDELITY INTERNATIONAL |
786,452 |
£10,145,231 |
| Hargreave Hale & Co |
784,942 |
£10,125,752 |
| Canaccord Genuity Group Inc |
780,008 |
£10,062,103 |
| Artemis Investment Management Ltd |
729,575 |
£9,411,518 |
| Norges Bank |
683,030 |
£8,811,087 |
| Charles Stanley |
659,277 |
£8,504,673 |
| Jonathan M Gumpel |
642,460 |
£8,287,734 |
| Brewin Dolphin Limited |
622,157 |
£8,025,825 |
| Chelverton Asset Management |
609,008 |
£7,856,203 |
| JTC plc |
603,261 |
£7,782,067 |
| Amati Global Investors |
572,459 |
£7,384,721 |
| Rathbones Investment Management |
488,191 |
£6,297,664 |
| Kames Capital |
487,376 |
£6,287,150 |
| Spearpoint Holdings Limited |
428,227 |
£5,524,128 |