| Aberforth Partners |
24,802,909 |
£34,972,102 |
| J O Hambro Capital Management Limited |
11,505,303 |
£16,222,477 |
| Perpetual Limited |
11,315,996 |
£15,955,554 |
| Huntington Management, LLC |
9,852,275 |
£13,891,708 |
| BlackRock Investment Managers (UK) |
9,291,081 |
£13,100,424 |
| Morgan Stanley |
8,325,023 |
£11,738,282 |
| The Wellcome Trust Limited |
7,615,301 |
£10,737,574 |
| ACR Alpine Capital Research |
6,871,037 |
£9,688,162 |
| Canaccord Genuity Group Inc |
6,361,026 |
£8,969,047 |
| 1607 Capital Partners, LLC |
5,775,000 |
£8,142,750 |
| Soros Fund Management |
5,602,726 |
£7,899,844 |
| Alantra EQMC Asset Management, SGIIC, S.A. |
5,413,717 |
£7,633,341 |
| Santander Asset Management UK |
5,267,323 |
£7,426,925 |
| Schroder Investment Management |
5,145,288 |
£7,254,856 |
| Columbia Threadneedle Investments |
5,020,929 |
£7,079,510 |
| 22NW, LP |
4,967,423 |
£7,004,066 |
| SFM UK Management LLP |
4,761,276 |
£6,713,399 |
| Chelverton Asset Management Limited |
4,678,875 |
£6,597,214 |
| Allianz Global Investors |
3,641,634 |
£5,134,704 |
| Premier Miton Investors |
3,633,000 |
£5,122,530 |
| Royal London Asset Management |
3,549,000 |
£5,004,090 |
| Bank of New York |
3,495,889 |
£4,929,204 |
| Henderson Global Investors |
3,375,404 |
£4,759,320 |